00:01
The vas deferens again
is very different. The ductus deferens or
vas deferens is a muscular tube. It has a
lot of smooth muscles wrapped around it. And
the layers of smooth muscle are orientated
in different directions, again, to ensure very
rapid force of all expulsion of the spermatozoa
in the lumen during ejaculation. The epithelium
also contains stereocilia, and sometimes the
epithelial forms this longitudinal fold, which
often might seem quite a characteristic structure
when you compare them with other ducts, and
not only in the male reproductive system,
but in other tubes of the body. Probably the
most distinguishing characteristic here though
is the very, very thick smooth muscle layers
around the wall. Here is a section through
the spermatic cord. The spermatic cord at
least contains a number of components.
01:14
On the left-hand side of this section, the tube
or the very small lumen and the thick muscular
wall is the vas deferens or ductus deferens.
Other structures you see here that dominate
are blood vessels, more towards the right-hand
edge of the section. Those blood vessels consist
of components or branches of the testicular
artery, and also, veins of the pampiniform
plexus. When I spoke about the testis in another
lecture, I explained that the testis has to
be maintained outside the body cavity. Spermatogenesis
only occurs if the temperature of the testis
is about two to three degrees below normal
body temperature. And that’s achieved because
the blood vessel supplying the testis, the
testicular artery, coils as it moves down towards
the testis, and it has a very intimate relationship
to the veins that drain the testis, the pampiniform
plexus of veins. And those veins carry cooler
blood back to the body, back to the heart.
02:38
And the relationship between the testicular
artery, being very coiled, and this pampiniform
plexus enables the blood to become to be cooled
by these veins as it passes down towards the
testis. That’s the primary role of this pampiniform
plexus. It’s the primary structural
specialization that cools the testis by cooling
the blood leading to the testis. You can’t
see it clearly on this slide, but if you look
very carefully around the vas deferens, the
rather red stained regions represent section
through the cremaster muscle. The cremaster
muscle lifts the testis or lowers the testis
in the scrotum. Scrotum also has a duct muscle
through it and that contracts or relaxes.
And this process of contracting or relaxing
creates the scrotum as being rather a tight
sac holding the testis or rather a loose flaccid sac,
and that’s designed to conserve heat loss
or create heat again in a cold temperature,
for instance, or heat retention in cold temperatures
that scrotum will contract and wrinkle to
maintain or at least reduce heat loss. So
you have this variety of mechanisms designed
to make sure the testis operates at a temperature
below body temperature.